The reliquary itself is collapsing into a much deeper dungeon. It can be entered from the jungle entrance in an old bullywug temple, or by climbing up out of the deeper levels below. But getting into the reliquary proper to acquire the holy relics within requires collecting the Agate, Jade, and Lapis keys that are sealed into tombs around the space.
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(Source: Dyson’s Dodecahedron)